早更新世希腊埃楚斯堪熊的新材料.pdf

Quaternary International xxx (2017) 1e18 Contents lists available at ScienceDirect Quaternary International journal homepage: /locate/quaint Revisiting Ursus etruscus (Carnivora, Mammalia) from the Early Pleistocene of Greece with description of new material George D. Koufos a, *, George E. Konidaris b, Katerina Harvati b a Aristotle University of Thessaloniki, Department of Geology, Laboratory of Geology and Palaeontology, 54124, Thessaloniki, Greece b Eberhard-Karls University of Tübingen, Palaeoanthropology, Senckenberg Centre for Human Evolution and Palaeoenvironment, Rümelinstr. 23, 72070, Tübingen, Germany a r t i c l e i n f o a b s t r a c t Article history: The Villafranchian ursids of Greece are scanty, represented by a few isolated teeth and bones. During our Received 9 May 2017 last field campaigns in the Mygdonia Basin (Macedonia, Greece) we discovered an almost complete Received in revised form cranium, as well as some cranial, dental and postcranial remains, which are presented in this article. The 20 September 2017 new material originates from the locality Tsiotra Vryssi (TSR), dated to the late Villafranchian (1.8e1.2 Accepted 28 September 2017 Available online xxx Ma). The specimens are described and compared with ursids from various European localities; their morphological characters and dimensions place it with Ursus etruscus Cuvier, 1823, while the dental