(植物生理课件）Chapter5同化物的运输与分配.ppt

Chapter 5 同化物的运输与分配p146-167 1. 同化物的运输 2. 同化物的装入与卸出 3. 同化物的配置与分配 4. 影响同化物运输与分配的外部因素 Section 1 Translocation systems of assimilate in plant Section 2 Translocation systems of assimilate in plant Fruit stores photoassimilate 二、同化物运输的途径 （一）胞内和胞间运输 1.胞内运输 2.胞间运输 质外体和共质体 数目多 106~107/mm2 in plant The plasmodesma is ?3nm, can transport the limited molecule of 800-1000D, but virus 10000D. Plasmodesma function： Substance transport and information transduction.物质运输与信息传递 Less resistance to substance transport cell by cell because of no transmembrane transport. 运输阻力小 Such as the resistance for plasma membrane is 0.31Ω/m2，for tonoplast 0.1Ω/m2 and for plasmadesma 0.05/Ωm2. 3. Short distance translocation is alternative between apoplast and symplast 3. 共质体和质外体交替运输 转移细胞的功能（源端装入，库端卸出） Transfer cells :A group of specific cells exist in the terminal in conduct tissue, flower or fruits 输导组织末端及花果器官 特化细胞 Characteristics in structure and metabolism : Large surface area of cell because cell wall and plasma membrane show rugocity (ingrowth). Abundant cytosol and organelles, high metabolic activity . Companion cells and intermediary cells are considered as transfer cells. Main function：Loading assimilate into phloem from source and unloading assimilate into sink cell from phloem or other conduct tissue. （二）器官间的运输 Section 3 Phloem transport of assimilates  3.1 Elements of phloem sap How to get phloem sap? Table 5-1 phloem sap from yucca Total dry matter 17.1-19.2(%) K 1.68 Amino acid (%) EC (ms/cm) 1.03 Mg 0.051 Gln 58 pH 8.0-8.2 Ca 0.014 Val 10 Components (mg/ml) Na 0.0014 Ser,Gly 7 Sucrose 150-165 Zn 0.0021 (Iso-)Leu 6 Glucose 2-4 Fe 0.0041 Lys 5 Fructose 2-4 Mn 0.0005 Glu 4 Protein 0.5-0.8