珊瑚树小孢子的发育MicrosporeDevelopmentofViburnum.PDF

32(1) 51-62, 2010 51 林業研究季刊 ： 研究報告 珊瑚樹小孢子的發育 1 2 3 張焜標 許秋容 張雯婷 【摘要】珊瑚樹花藥壁層的發育屬於雙子葉型，花藥內壁層於花粉成熟時細胞壁呈徑向加厚。營養 層則為分泌型，且由單核之細胞所組成。小孢子之發育可分為雄蕊始原期、小孢子母細胞期、減數 7 分裂期、四分體期、液胞化與自由小孢子期、雙細胞花粉粒期和花粉成熟與開花期等 個階段。小 孢子母細胞減數分裂後行同時型之胞質分裂，產生之四分體呈四面體形，外圍由胼胝質圍繞。成熟 之花粉粒具三溝孔，且表面為不規則之網紋。 【關鍵詞】珊瑚樹、胼胝質、花粉粒、營養層、四分體 Research paper Microspore Development of Viburnum odoratissimum Ker Kun-Piao Chang1 Chiou-Rong Sheue2 Wen-Ting Chang3 Abstract The development of anther wall layers in Viburnum odoratissimum belonged to 【 】 dicotyledonous type and the cell wall endothecial layer showed radial thickening during pollen maturation. The tapetum was a secretory type and composed of single nucleate cells. The development of microspores could be divided into seven stages that were stamen primordial stage, microspore mother cell stage, meiosis stage, tetrad stage, vacuolated and free microspore stage, two-cellular pollen grain stage, and pollen maturation and anthesis stage. Microspore mother cells undertook meiosis with cytokinesis simultaneously to produce tetrahedral tetrads enclosed within callose wall. The surface of mature pollen grains was tricoporate with irregular reticulate sculptures. Key words Callose, pollen grain, tapetum, tetrad, Viburnum odoratissimum. 【 】 1. 912 1 國立屏東科技大學森林學系教授， 屏東縣內埔鄉學府路 號。 Professor, Department of Forestry, National Pingtung University of Science and Technology, 1 Hseuh Fu Rd., Neipu 912, Pingtung Taiwan. 2 402 250 國立中興大