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BULLETIN OF
MATHEMATICAL BIOPHYSICS
VOLUME 5, 1943
A L O G I C A L C A L C U L U S O F T H E
I D E A S I M M A N E N T I N N E R V O U S A C T I V I T Y
WARREN S. MCCULLOCH AND WALTER PITTS
FROM THE UNIVERSITY OF ILLINOIS, COLLEGEOF MEDICINI~,
DEPARTMENT OF PSYCHIATRY AT THE ILLINOIS NEUROPSYCHIATRIC INSTITUTE,
AND THE UNIVERSITY OF CHICAGO
Because of the all-or-none character of nervous activity, neural
events and the relations among them can be treated by means of propo-
sitional logic. It is found that the behavior of every net can be described
in these terms, with the addition of more complicated logical means for
nets containing circles; and that for any logical expression satisfying
certain conditions, one can find a net behaving in the fashion it describes.
It is shown that many particular choices among possible neurophysiologi-
cal assumptions are equivalent, in the sense that for every net behav-
ing under one assumption, there exists another net which behaves un-
der the other and gives the same results, although perhaps not in the
same time. Various applications of the calculus are discussed.
I. Introduction
T h e o r e t i c a l n e u r o p h y s i o l o g y rests on c e r t a i n c a r d i n a l a s s u m p -
tions. T h e n e r v o u s s y s t e m is a n e t of neurons, each h a v i n g a s o m a
and a n axon. T h e i r a d j
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