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4 Update TRENDS in Ecology and Evolution Vol.20 No.1 January 2005
The evolution of large size: how does Cope’s Rule
work?
David W.E. Hone and Michael J. Benton
Department of Earth Sciences, Wills Memorial Building, Queens Road, Bristol, UK, BS8 1RJ
Cope’s Rule is the tendency for organisms in evolving Size and fitness
lineages to increase in size over time. The concept is The results are quite startling (Figure 1): size increase
detailed in many textbooks, but has rarely been demon-
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