adaptive radiation of gobies in the interstitial habitats of gravel beaches accompanied by body elongation and excessive vertebral segmentation适应辐射间隙鰕虎鱼的栖息地的砾石海滩伴随着身体伸长和脊椎过度分割.pdfVIP

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adaptive radiation of gobies in the interstitial habitats of gravel beaches accompanied by body elongation and excessive vertebral segmentation适应辐射间隙鰕虎鱼的栖息地的砾石海滩伴随着身体伸长和脊椎过度分割.pdf

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adaptive radiation of gobies in the interstitial habitats of gravel beaches accompanied by body elongation and excessive vertebral segmentation适应辐射间隙鰕虎鱼的栖息地的砾石海滩伴随着身体伸长和脊椎过度分割

BMC Evolutionary Biology BioMed Central Research article Open Access Adaptive radiation of gobies in the interstitial habitats of gravel beaches accompanied by body elongation and excessive vertebral segmentation Tomohiko Yamada*, Tomoshige Sugiyama, Nana Tamaki, Atsushi Kawakita and Makoto Kato Address: Graduate School of Human and Environmental Studies, Kyoto University, Yoshida-Nihonmatsu-cho, Sakyo, Kyoto 606-8501, Japan Email: Tomohiko Yamada* - luciogobius@star.ocn.ne.jp; Tomoshige Sugiyama - sugiyama.t@; Nana Tamaki - egocoro@; Atsushi Kawakita - kawakita@s01.mbox.media.kyoto-u.ac.jp; Makoto Kato - kato@zoo.zool.kyoto- u.ac.jp * Corresponding author Published: 28 June 2009 Received: 13 March 2009 Accepted: 28 June 2009 BMC Evolutionary Biology 2009, 9:145 doi:10.1186/1471-2148-9-145 This article is available from: /1471-2148/9/145 © 2009 Yamada et al; licensee BioMed Central Ltd. This is an Open Access article distributed under the terms of the Creative Commons Attribution License (/licenses/by/2.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited. Abstract Background: The seacoasts of the Japanese Arc are fringed by many gravel beaches owing to active tectonic uplift and intense denudation caused by heavy rainfall. These gravel beaches are inhabited by gobies of the genus Luciogobius that burrow into the gravel sediment and live interstitially. Although their habitat and morphology (e. g., reduced fins, el