A New Leptoceratopsid (Ornithischia Ceratopsia) from the Upper Cretaceous of Shandong, China and Its Implications for Neoceratopsian Evolution 英文参考文献.docVIP

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A New Leptoceratopsid (Ornithischia Ceratopsia) from the Upper Cretaceous of Shandong, China and Its Implications for Neoceratopsian Evolution 英文参考文献.doc

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A New Leptoceratopsid (Ornithischia Ceratopsia) from the Upper Cretaceous of Shandong, China and Its Implications for Neoceratopsian Evolution 英文参考文献

ANewLeptoceratopsid(Ornithischia:Ceratopsia)from theUpperCretaceousofShandong,ChinaandIts ImplicationsforNeoceratopsianEvolution XingXu1*,KebaiWang2,XijinZhao1,CorwinSullivan1,ShuqingChen2 1KeyLaboratoryofEvolutionarySystematicsofVertebrates,InstituteofVertebratePaleontologyandPaleoanthropology,ChineseAcademyofSciences,Beijing,China, 2ZhuchengDinosaurMuseum,BureauofTourism,Zhucheng,Shandong,China Abstract Background:Theceratopsiansrepresentoneofthelastdinosaurianradiations.Traditionallytheonlyuniversallyaccepted speciosecladewithinthegroupwastheCeratopsidae.However,recentdiscoveriesandphylogeneticanalyseshaveledto therecognitionofanewspecioseclade,theLeptoceratopsidae,whichispredominantlyknownfromtheUpperCretaceous ofNorthAmerica. Methodology/Principal Findings: Here we report a new leptoceratopsid taxon, Zhuchengceratops inexpectus gen. et sp. nov.,basedonapartial,articulatedskeletonrecoveredfromtheUpperCretaceousWangshiGroupofZhucheng,Shandong Province,China.AlthoughZhuchengceratopsissignificantlydifferentfromotherknownleptoceratopsids,itisrecoveredasa derived member of the group by our phylogenetic analysis. Furthermore, Zhuchengceratops exhibits several features previouslyunknowninleptoceratopsidsbutseeninceratopsidsandtheircloserelatives,suggestingthatthedistributionof morphologicalfeatureswithinceratopsiansismorecomplexthanpreviouslyrealized. Conclusion/Significance:ThediscoveryofZhuchengceratopsincreasesboththetaxonomicdiversityandthemorphological disparity of the Leptoceratopsidae, providing further support for the hypothesis that this clade represents a successful radiationofhorneddinosaursinparallelwiththeCeratopsidaeintheLateCretaceous.Thisdocumentsasurprisingcaseof the coexistence and radiation of two closely-related lineages with contrasting suites of jaw and dental features that probablyreflectadaptationtodifferentfoodresources. Citation:XuX,WangK,ZhaoX,SullivanC,ChenS(2010)ANewLeptoceratopsid(Ornithischia:Ceratopsia)fromtheUpperCretaceousofShandong,