黄颡鱼性腺发育及周年变化的研究-水产养殖专业毕业论文.docxVIP

摘 摘 要 黄颡鱼Pesudobugrusfulvidraco(Richardson)性细胞发育按其组织学特点，可 分为6个时相，性腺依其形态结构特征，可分为6个时期。黄颡鱼卵巢～对，长 囊状。30～60d龄的鱼处于第1期卵巢，且终身只出现一次，主要由卵原细胞构 成，外周无滤泡细胞，核内有一个明显的核仁；60～270d龄及每年9月～次年3 月性成熟的鱼处于第1I期卵巢，II时相卵母细胞出现明显的卵黄核；270～300d 龄及每年3～4月份的性成熟的鱼处于第1II期卵巢，In时相卵母细胞出现放射膜 和带状细丝：310d龄的鱼及每年5月份左右性成熟的鱼处于第Ⅳ期卵巢，Ⅳ时 相卵母细胞的受精孔和精孔细胞明显，核出现极性；330d龄及每年6~7月份性 成熟的鱼处于第V期卵巢，第V时相卵母细胞中卵黄颗粒融合现象明显。黄颡鱼 精巢一对，分生精部与贮精囊。30～60d龄的幼鱼处于1期精巢，精原细胞分散 于间质细胞中；60～130d龄及每年10月一次年4月性成熟的鱼处于II期；90日 龄以上的鱼，精巢发育至In期，成熟系数可达O．32％～0．66％。精子的发生经历 精原细胞、精母细胞、精子细胞及成熟的精子等阶段，精子无顶体，核凹窝，精 子尾部细长，轴丝横切面为典型的“9+2”结构。 黄颡鱼在池塘饲养和投喂人工饲料条件下，其性成熟年龄为14个月，比同 地区天然水域自然群体(性成熟时间约23个月)早熟1年。繁殖季节为每年的 6～7月，属一次产卵类型。首次达性成熟年龄群体的体重为70～130克，最大 成熟系数达24％，绝对怀卵量3400～4500粒／尾，相对怀卵量50～60。 关键词： 黄颡鱼； 卵巢发育：精巢发育：形态组织；周年变化 Study Study on the development and Annual Change of Gonad in Pesudobugrus fulvidraco Abstract：According to the morphological and histological features，the development of gland can be divided into six phases．The ovaries of Pseudobagursfulvidraco can be divided into two cystiform parts．The ovaries are in stage 1 when the fishes are 30 to 60 days old and only appears once in their life．It is main to be constituted by oogenium，and the pefi【phery have no the follicle，nucleus inside to have all obvious of nucleolus；when they are 60～270 days old and every year September to next year the fish of mature Mar．，the ovaries began to grow to stage II the yolk nucleus are very distinct；When they are 270～300 days old and the fish of sex mature of every year Mar．to Apri．，the ovaries enter into stage III，radiation theca and fillet filament cart be seen in the stage III；The ovaries grow to stage IV when they are 3 1 0 days old and every year of May．the fish ofmature or so,the micropyle and the micropylar cell are very clear,nucleus emergences polarity．After 330 days old and every year June to July the fish for maturely,the ovaries enter into stage V，the fuse of yolk granule are clear． The spermaries ofPseudobagursfulvidraco can be divided into two parts．The anterior part is s