棘龙科方形形态功能分析（恐龙：兽脚亚目）和棘背龙的存在和在北非的赛诺曼第二spinosaurine分类.pdfVIP

RESEARCH ARTICLE Morphofunctional Analysis of the Quadrate of Spinosauridae (Dinosauria: Theropoda) and the Presence of Spinosaurus and a Second Spinosaurine Taxon in the Cenomanian of North Africa. Christophe Hendrickx1,2¤ *, Octávio Mateus1,2, Eric Buffetaut3 1 GeoBioTec, Faculdade de Ciências e Tecnologia, Universidade Nova de Lisboa, Caparica, Portugal, 2 Museu da Lourinhã, Lourinhã, Portugal, 3 CNRS (UMR 8538), Laboratoire de Géologie de l’École Normale Supérieure, PSL Research University, Paris, France ¤ Current address: Evolutionary Studies Institute, Center of Excellence in Palaeosciences, University of the Witwatersrand, Johannesburg, South Africa * christophendrickx@ OPEN ACCESS Citation: Hendrickx C, Mateus O, Buffetaut E (2016) Abstract Morphofunctional Analysis of the Quadrate of Spinosauridae (Dinosauria: Theropoda) and the Presence of Spinosaurus and a Second Spinosaurine Six quadrate bones, of which two almost certainly come from the Kem Kem beds (Cenoma- Taxon in the Cenomanian of North Africa. PLoS ONE nian, Upper Cretaceous) of south-eastern Morocco, are determined to be from juvenile and 11(1): e0144695. doi:10.1371/journal.pone.0144695 adult individuals of Spinosaurinae based on phylogenetic, geometric morphometric, and Editor: Al