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prediction of breeding values when variances are not known育种值预测方差是未知的
Prediction of breeding values when variances are not known
D. GIANOLA* J.L. FOULLEY* R.L. FERNANDO’
* Department of Animal Sciences
University of Illinois at (lrbana Champaign, U.S.A.
**
LN.R.A., Station de Génétique quantitative et appliquee
Centre de Recherches Zootechniques, F 78350 Jouy-en-Josas
Summary
The joint distribution of breeding values and of records usually depends on unknown
parameters such as means, variances and covariances in the case of the multivariate normal
distribution. If the objective of the analysis is to make selection decisions, these parameters should
be considered as « nuisances ». If the values of the parameters are unknown, the state of
uncertainty can be represented by a prior probability distribution. This is then combined with the
information contributed by the data to form a posterior distribution from which the needed
predictors are calculated after integrating out the « nuisances ». Prediction under alternative states
of knowledge is discussed in this paper and the corresponding solutions presented. It is shown that
when the dispersion structure is unknown, modal estimators of variance parameters should be
considered. Because a Bayesian framework is adopted, the estimates so obtained are necessarily
If about means and variances is and the distribu-
non-negative. prior knowledge completely vague
tion is multivariate normal, the « optimalpredictors in the sense of maximizing the expected
merit of the selected candidates can be approximated by using the « mixed model equations » with
the unknown variances replaced by restricted maximum like
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