短柄五加大小孢子发生和雌雄配子体发育的研究-植物研究.PDF

　　　　　　　　　 BULLETIN OF BOTANICAL RESEARCH 18 　2 1998 　4 Vol.18　No.2 April, 　1998 、、 1 1 2 1 王仲礼 　刘林德 　田国伟 　申家恒 (1., 264025)(2., 100871) 　　短柄五加花药5 枚, 每 个花药四个花粉囊。小孢子母细胞减数分裂时, 胞 质分裂为同时型, 产生正四面 形的四分 。 花药壁由表皮、药室内壁、中层和绒 毡层四层细胞组成, 其发育类型为双子叶型。腺质绒毡层, 其细胞为二核。三细胞 型花粉。子房5 室, 每室两 个胚珠, 上胚珠败育, 下胚珠可育。下胚珠倒生, 具单珠 被, 厚珠心。大孢子母细胞减数分裂形成线性排列的四个大孢子, 雌配子 发育属 蓼型。开花当天, 花粉散开, 雌配子 尚未成熟, 处于三 个反足细胞退化的四细胞 胚囊阶段。短柄五加为雄蕊先熟植物。记述了雌、雄蕊发育的对应关系。 　短柄五加;小孢子;雄配子 ;大孢子;雌配子 ;雄蕊先熟 STUDIES ON THE MEGASPOROGENESIS AND MICROSPOROGENESIS AND THE DEVELOPMENT OF THEIR FEMALE AND MALE GAMETOPHYTE IN ELE UTHER OCOCCUS BRA CH YP US 1 1 2 1 Wa g Zho g-li 　Liu Li -de 　Tia Guo-wei 　She Jia-he g (1.Departme t of Biology, Ya tai Normal U iversity, Ya tai 264025) (2.College of Life scie ces, Peki g U iversity, Beiji g 100871) Abstract 　The flow er of E leutherococcus br achyp us has five a thers, each a ther comprises four microspora gia.I meiosis of microsporocyte, cytoki esis is simulta e- ous type, which produces a isobilateral tetrad.A ther w all i cludes epidermis, e - dothecium, middle layer a d tapetum, which belo gs to dicotyledo ous type.Tape- tum with bi ucleate cells is of the secretory type.The mature polle grai is tricellu- ar.The ovary is five-loculus w ith two ovules i each.The upper ovule is sterile a d the lower ovule is fertile.The latter is mo o-i tegi ous, crassi ucellate a d a otro- (NO 　:1997-9-1 178