植物逆境信息传递研究进展.ppt

Stress Signaling in Plants 贾文锁 中国农业大学;2;3;4;一 “感知”和“反应” ——生物生存和发展的基础 ;6;7;敏感植物;9;10;11; 一般植物;13;14;Rain,Wind, and Touch-Induced Expression of Calmodulin and Calmodulin-Related Genes in ArabidopsisCell 60,357-364,1990;16;二;1.形态、解剖;Cluster Roots;2.整体生理;21;生命周期—短命菊;3.细胞生理及生化;渗透调节;保护酶系统与自由基清楚剂 游离的带有不成对电子的分???、原子或离子;26;27;区隔化机制 ;结构避损机制;功能蛋白 代谢调控蛋白 （渗透调控系统，保护酶） Lea 蛋白（Late-Embryogenesis-Abundant Protein） 通道蛋白：离子通道，水通道 热击蛋白：亲水，保护酶活 损伤修复蛋白 调节蛋白 信号蛋白 转录因子 ;31;基因;33;应答反应与伤害反应;三; Cellular Singling System ?;;细胞信号系统;1 Receptor /Sensor 2. Second Messenger 3. Protein Kinase 4. Protein Phosphatase 5. G-protein 6. Transcription Factor 7. Ubiquitination 8. Methylation ;(1) Receptor /Sensor?;41;42;43;Receptor tyrosine kinase; 细胞内受体;Examples of Phytohormone Recepor and Signal Transduction;47;ABA Receptor ;49;BRI1 is a membrane-associated receptor that cycles between the plasma membrane and endosomal compartments. The extracellular leucine-rich repeat domain binds brassinosteroids and transduces the signal through an intracellular kinase domain. GTG1 and GTG2 are GPCR-type G proteins that bind abscisic acid. They have inherent GTPase activity but also interact with the only canonical G subunit in Arabidopsis. PYR1/RCAR1 is a soluble ABA receptor that represses PP2C phosphatases in the presence of ABA. The cytokinin receptors CRE1, AHK2 and AHK3 are plasma-membrane-associated and perceive cytokinin through their extracellular domains. Cytokinin binding triggers a phosphorylation cascade that is ultimately transmitted to response regulators in the nucleus. Like the cytokinin receptors, the known ethylene receptors are two-component regulators. All five receptors are active in the endoplasmic reticulum and transmit their signal through a common downstream component called CTR1. TIR1 and COI1 are F-box pro