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2023年托福阅读辅导:老托福阅读真题及答案
passage11整理
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老托福阅读文本passage11
Plantsaresubjecttoattackandinfectionbyaremarkablevarietyof
symbioticspeciesandhaveevolvedadiversearrayofmechanisms
designedtofrustratethepotentialcolonists.Thesecanbedividedinto
preformedorpassivedefensemechanismsandinducibleoractive
systems.Passiveplantdefensecomprisesphysicalandchemicalbarriers
thatprevententryofpathogens,suchasbacteria,orrendertissues
unpalatableortoxictotheinvader.Theexternalsurfacesofplants,in
additiontobeingcoveredbyanepidermisandawaxycuticle,oftencarry
spikyhairsknownastrichomes,whicheitherpreventfeedingbyinsects
ormayevenpunctureandkillinsectlarvae.Othertrichomesaresticky
andglandularandeffectivelytrapandimmobilizeinsects.
Ifthephysicalbarriersoftheplantarebreached,thenpreformed
chemicalsmayinhibitorkilltheintruder,andplanttissuescontaina
diversearrayoftoxicorpotentiallytoxicsubstances,suchasresins,
tannins,glycosides,andalkaloids,manyofwhicharehighlyeffective
deterrentstoinsectsthatfeedonplants.ThesuccessoftheColorado
beetleininfestingpotatoes,forexample,seemstobecorrelatedwithits
hightolerancetoalkaloidsthatnormallyrepelpotentialpests.Other
possiblechemicaldefenses,whilenotdirectlytoxictotheparasite,may
inhibitsomeessentialstepintheestablishmentofaparasiticrelationship.
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Forexample,glycoproteinsinplantcellwallsmayinactivateenzymes
thatdegradecellwalls.Theseenzymesareoftenproducedbybacteria
andfungi.
Activeplantdefensemechanismsarecomparabletotheimmune
systemofvertebrateanimals,althoughthecellularandmolecularbases
are
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