Buxus Leaves from the Oligocene of Guangxi, China and Their Biogeographical Significance.pdfVIP

Vo1．89No．5PP．1453—1469 ACTAGEOLOGICASINICA (EnglishEdition) Oct．2015 BuxusLeavesfrom theOligoceneofGuangxi，ChinaandTheir BBli00ggee00ggrraapphnliccaallSSiignniifliiccaannccee MAFujun，，。一，WANGQiujun，DONGJunling，YANGYi，WANGWenjia， YAN DefeiandSUN Bainian，，， lSchoolofEarthSciences＆KeyLaboratoryofMineralResourcesinWesternChina(GansuProvince)． La nzhouUniversi砷，Lanzhou73oooo,China 2KeyLaboratoryofPetroleumResources，GansuProvince．CA Lanzhou37oooo,China 3SchoolofGeographyandTourism，GuizhouNormalCollege，Guiyang550018,China 4KeyLaboratoryofEconomicStratigraphyandPalaeogeography,NanjingInstituteofGeologyand Palaeontology．CA ．Nanjing210008,China． Abstract：The presentpaper reports two new species ofBuxus from the Oligocene Ningming Formation ofGuangxi，South China，on the basis of severalcompressed fossilleaves．BUXUS ningmingensissp．nov．ischaracterizedbyanelliptictoovateshape；pinnatevenationwith marginal secondary，longhairsontheadaxialsideofthemidveinbase；andamphistomatic。anomocyticstoma~ withaprom inentouterring．A11ofthesecharacteristicssuggestaclosestaffinitytothelivingBuxus microphyllasubsp．sinica．Anothernew speciesidentifledasBuxuspreaustro-yunnanensissp．nov．is characterized bythediagnostic~rmationofits admediallybranchedtertiaries，thetrunksofwhich areoften stronglythickened bysclereides．ThisspeciesismostsimilartotheextantBuxu$austro． yunnanensis．Theco-occurrenceofBUXUSningmingensissp．nov．andBuxuspreaustro-yunnanensissp． nov．suggests thatthesetwoBUXUSlivedduringtheOligoceneunderawarm ，tropical tosubtropical climate，sim ilartothecurrentclimateofNingming．Onrfindingsprovidetheearliestfossilevidenceof Buxusleaveswithdetailedillustrationsofleafarchitecturalandcuticularfeatures．Theoc