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THEORETICAL POPULATION BIOLOGY 28, 1-17 (1985)
Population Dynamics in Variable Environments.
VI. Cyclical Environments*
SHRIPAD TULJAPURKAR
PhvsicFjEnvironmentaI Science,t Portland State University, Portland, Oregon 97207,
and Graduate Group in Demography, University of California,
2234 Piedmvnt Avenue, Berkeley, Cal@ornia 94720
Received December 7, 1983
This paper studies the dynamics of an age-structured population which experien-
ces cyclical variation in vital rates. The principal features of population behavior
are found to be contained in an explicitly calculable response function. Three dis-
tinct regimes of qualitative behavior are described when cycle period is respectively
much less than, of the order of, and much greater than the average generation
length. These results make explicit the way in which transient properties
corresponding to average vital rates determine population response to cycles.
‘1” 19X5 Academic Press, Inc
1. INTRODUCTION
MacArthur (1968) studied a population with two age classessubject to
cycles in juvenile survival and found population response to cycles to be
biologically significant and mathematically subtle. While his results clearly
warrant generalization his analysis does not generalize,
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