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Competitive Exclusion in a Discrete-time,
Size-structured Chemostat Model
Hal L. Smith
Department of Mathematics
Arizona State University
Tempe, AZ 85287–1804, USA
E-mail: halsmith@asu.edu
Xiao-Qiang Zhao
Department of Mathematics and Statistics
Memorial University of Newfoundland
St. John’s, NF A1C 5S7, Canada
E-mail: xzhao@math.mun.ca
May 22, 2000
Abstract Competitive exclusion is proved for a discrete-time, size-structured, nonlinear
matrix model of -species competition in the chemostat. The winner is the population
able to grow at the lowest nutrient concentration. This extends the results of earlier
work of the first author [11] where the case = 2 was treated.
Key words and phrases: Chemostat, discrete-time, size-structured model, competi-
tive exclusion.
AMS subject classifications: 34C35, 92A15
1 Introduction
The classical chemostat model of microbial growth and competition for a limiting sub-
strate has played a central role in population biology. See [12] for a treatment of chemo-
stat models. However, the classical model ignores the size structure of the population
Supported by NSF Grant DMS 9700910.
Supported in part by the NSERC of Canada.
1
and the observation that many microbes roughly double in size before dividing. Size-
structured chemostat models formulated by Metz and Diekmann [10] and by Cushing
[3, 4] (see also [12]) lead to hyperbolic partial differential equations with nonlocal bound-
ary conditions. A conceptually simpler approach to modeling size structure was taken
by Gage, Williams and Horton in [5
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