金粟兰AP1CHL基因功能的研究-生物学·植物学专业论文.docxVIP

苎塑堑茎查兰堡主兰竺堡苎Abstract 苎塑堑茎查兰堡主兰竺堡苎 Abstract Flowering is an important stage in the transition from vegetative growth to reproductive growth in the lives ofplants．The difierentiation and development offloral organs have been the foCUS of biologists’study,not only for也e variety of floral shape and other characteristics，but also for its important economic value． Two classes ofgenes control the development offlower：flower meristem identity genes which promote the initiation of flower meristem from iI迁k坩e90ence meristem．and the floral organ identity genes which，also named homeotic genes。determine which kind of floral organs will initiate．The flower meristem identity genes can activate the homeotic genes to sc}me extent． MADS．box genes family plays a key role in the molecular regulation of floral development，and most of them in plants belong to the typelI class，that is’the gene sequence is composed of M，I，K and C domains．The M domain,which encodes the DNA-binding domain and plays the rolesofboth binding DNA and forming dimmers．is the most conserved region．According to the classical“J蛆C”model．three classes of gelies act to speci母floral organs，namely sepals(A only)，petals(A十B)stamens(B+C)or carpels(C only)．Later,some new found MADS．box genes result in the“ABCD”model．“ABCDE” model and“quartet’model．All of these models make the understanding of floral ‘ development deeply． Chloranth搬spicatus belongs to Chloranthaceae Chloranthus according to its phylogenic position．It has no perianth and is one ofthe oldest angiosperm that have fossil． In the classification ofangiosperm，the simplism offloral organ is thought to be a derivative identity．But some new data about fossil show that the small and simple flowers appeared before the eomplicated flowers,like the flower of Chloranthus spicatus．Thus Chloranthus spicatus’special morphological characteristics and position in the plant Phylogenic tree ptays a great role in the research ofthe origin ofperianth and flowers． APIC舭gene is clone