光可以间接影响Rubisco的活性.PPT

生化Ch21 partIII重點整理 生科2A 0993B013許嘉珊 0993B035張以潔 心智圖 Photorespiration 當氧氣過剩、二氧化碳不足時，植物行Photorespiration(光呼吸作用) Rubisco具兩種催化能力：carboxylase oxygenase，當二氧化碳不足時oxygenase活性上升，分解RuBP RuBP + O2→ 3-PG + Phosphoglycerate 　3-PG可進入糖解作用，phosophoglycerate可被代謝成glycine，均能有效利用，但RuBP減少會降低光合作用的效率 Hatch-Slack pathway 目的：傳遞CO2至其他細胞 參與酵素：Pyruvate-Pi dikinase 　　　　　　　　　　　 　　　　　　(ATP→AMP+2Pi) 　　　　　　 Pyruvate-Pi dikinase 反應機構： 　　E-His + AMP-P-P +Pi → E-His-P + AMP + PPi 　　E-His-P + pyruvate → PEP + E-His 　Net : ATP + Pi + pyruvate AMP + PEP + PPi 調控：Threonine phosphorylation→抑制 C4 plant C3植物行一般的固碳途徑 C4植物行Hatch-Slack pathway 　能夠有效的運用光能 Question 2 Which statement about photosynthesis in C4 plants is NOT true? (A) the first product of carbon dioxide fixation is a compound with 4 carbon atoms. (B) C4 photosynthesis is an adaptation for plants living in hot, arid climates. (C) carbon dioxide is initially fixed in mesophyll cells, but the Calvin cycle is active in bundle sheath cells in leaves of C4 plants. (D) less ATP is used overall for sugar biosynthesis in C4 than in C3 plantsC4 photosynthesis is an adaptation for plants living in hot, arid climates like Tucson, Arizona. C4 plants initially fix CO2?in mesophyll cells as 4-carbon compounds, and later release CO2?in bundle sheath cells. There is an additional ATP requirement for each carbon dioxide utilized in this pathway. (E) photorespiration is minimal in C4 plants compared to C3 plants. The fixation of carbon dioxide Rubisco regulation Dark reaction Calvin-Benson cycle (Ch.22) Photorespiration Hatch-Slack pathway C4 plant Pyruvate-Pi dikinase The fixation of carbon dioxide 5 C sugar + 1 C → 6 C intermediate 6 C intermediate → 2* 3 C sugar 5 C sugar: Ribulose-1,5-bisphosphate 1 C : carbon dioxide 6 C sugar : 2-carboxy-3-ketoarabinitol 1,5-bisphosphate 3 C sugar:3-phosphoglycerate /2007/07/fixing-carbon-rubisco-reaction.html The fixation of carbon dioxide Regulation of rubisco 1.將rubisco 變成active form: carbamylation + Mg2+ *