植物生理课件5Chapter05 translocation of photoassimilates知识介绍.pptVIP

Chapter 5 同化物的运输与分配p146-167;Section 1 Translocation systems of assimilate in plant;Section 2 Translocation systems of assimilate in plant;Fruit stores photoassimilate;二、同化物运输的途径 （一）胞内和胞间运输 1.胞内运输 2.胞间运输 质外体和共质体;Figure5-3 shows conceptual model for the cell-to-cell trafficking of viral RNA(vRNA) in uninfected plants.;Figure 5-4 shows a conceptual model for cell-to-cell trafficking of specific proteins of a size larger than passive SEL.;Plasmodesma function：;3. Short distance translocation is alternative between apoplast and symplast 3. 共质体和质外体交替运输;转移细胞的功能（源端装入，库端卸出） ;Transfer cells :A group of specific cells exist in the terminal in conduct tissue, flower or fruits 输导组织末端及花果器官 特化细胞 Characteristics in structure and metabolism : Large surface area of cell because cell wall and plasma membrane show rugocity (ingrowth). Abundant cytosol and organelles, high metabolic activity . ;Companion cells and intermediary cells are considered as transfer cells. Main function：Loading assimilate into phloem from source and unloading assimilate into sink cell from phloem or other conduct tissue.;Figure 5-10 Sieve elements and companion cells;（二）器官间的运输 ;Figure 5-11 conduct tissue;1. Sieve tube and companion cell（SE/CC）are in angiosperm;Figure 5-13 Photograph of sieve element and companion cell by electron microscope;P- protein：与植物各类和筛管分子的成熟度有关. 幼嫩筛管：扭曲盘绕的球形或纺锤形 成熟筛管：管状或丝状的结构 功能：促进物质运输；防止汁液的流失。 寿命：有长（2-10），有短（一个生长季节）;Section 3 Phloem transport of assimilates ;3.1 Elements of phloem sap How to get phloem sap?;Figure 5-15 harvest phloem sap;;Table 5-1 phloem sap from yucca ;Components in phloem sap: 1. Sugars. Sucrose 90%，a small number of sorbitol, mannitol and oligose including raffinose and stachyose. Main sorbitol in rose plant phloem. ;;2. Amino acids and Amides. Glu or Asp and their amides. ;3.proteins(enzymes, ATPase) and nuclear acid (small mRNA). 4. Plant hormones, ATP, glucolipid ,vitamins. 5.organic acids. Mal. ;6. inorganic ions：cationsanions，most K+ for cations, most P