第五节 植物的光形态建成.pptx

第五节 植物的光形态建成自然界中的光500700600400Wavelength(nanometers)植物的光形态建成（photomorphogenesis）Light-grownArabidopsisEtiolated characteristicsDe-etiolated characteristicsDistinct apical hook (dicot) or coleoptile (monocot) No leaf growth No chlorophyll Rapid stem elongation Limited radial expansion of stem Limited root elongation Limited production of lateral rootsApical hook opens or coleoptile splits open Leaf growth promoted Chlorophyll produced Stem elongation suppressed Radial expansion of stem Root elongation promoted Lateral root development accelerated Light ControlsPhotosynthesis Rate of Growth Direction of Growth Pigmentation Flowering Fruiting Leaf Fall Onset of Dormancy光敏素向光蛋白隐花色素Phototropins 向光素拟南芥的phot突变体拟南芥的phot突变体PHOT1 and PHOT2 mediate blue light-dependent stomatal openingKinoshita et al. 2001 Nature: 414, 656Phototropin proteins contain two LOV (Light, Oxygen, Voltage sensitive) domains and a downstream serine/threonine kinase. The chromophore is a Flavin mononucleotide (FMN) which is non-covalently bound within each LOV domain. Schematic illustration of the LOV and kinase domains of phototropin in light-induced phosphorylation. (a) In the dark, the phosphorylation activity of the kinase domain of phototropin is inhibited by the binding of the LOV2 domain. (b) Under light conditions, the phosphorylation activity of the kinase is induced by the release of the LOV2 domain./science?_ob=PublicationURL_tockey= /science?_ob=PublicationURL_tockey=Volume 9, Issue 5, October 2006, Pages 503-508 Cryptochromes隐花色素/Research/cryptochrome/Cryptochrome and the avian compassThe HY4 gene may define a blue light photoreceptor CRYPTOCHROME 1 (CRY1)Phenotypes of cry2 mutantcyr2 is late flowering under long day conditionsThe structure of cryptochromes Cryptochrome controlled deetiolation. In Arabidopsis, cry1 and cry2 regulate most of the blue-light-specific developmental programs. Shown are only phenotypic effects during deetiolation caused by mutations in cry1 or cry2. Arabidopsis